| D.Sc. (Medical Sciences) | Facultad de Medicina, Universidad de la República, Montevideo, URUGUAY | 1961 |
| B.Sc. (Biological Sciences) | Facultad de Medicina, Universidad de la República, Montevideo, URUGUAY | 1950 |
FELLOWSHIP
| US. Public Health Service, N.I.H. Fellow | 1961–1963 |
| Rockefeller Foundation Fellow | 1959–1961 |
POSTGRADUATE EDUCATION
| Research Associate | Biochemistry, Cornell University Medical College, New York, New York, USA | 1963–1965 |
| Fellow (N.I.H.) | Biochemistry, Cornell University Medical College, New York, New York, USA | 1961–1963 |
